Taxonomy and field experience
Gleason received his doctorate at Columbia in 1906, taught in Illinois and Michigan, and studied vegetation in the Philippines, Java and Ceylon. He worked at the New York Botanical Garden from 1918 to retirement in 1950, including research on South American plants and Melastomataceae.
References: [1]
Dispersal and establishment
His 1926 paper examined how seeds reach locations and establish under suitable conditions. Species differ in dispersal and environmental requirements; their overlapping distributions help produce the vegetation observed at a particular site.
References: [2]
Variable community boundaries
Abrupt environmental changes can produce distinct vegetation boundaries, while gradual changes may produce transitions. Repeated combinations of plants therefore need not imply fixed units with perfectly stable composition.
References: [2]
Not an absence of interactions
Gleason allowed that vegetation modifies the environment and that plants influence one another. His argument opposed treating an association as an organism; it did not claim that competition or other interactions never occur.
References: [2]